Wardhaugh, Carl W., Edwards, Will, and Stork, Nigel E. (2015) The specialization and structure of antagonistic and mutualistic networks of beetles on rainforest canopy trees. Biological Journal of the Linnean Society, 114 (2). pp. 287-295.

Wardhaugh, Carl W. (2014) The spatial and temporal distributions of arthropods in forest canopies: uniting disparate patterns with hypotheses for specialisation. Biological Reviews, 89 (4). pp. 1021-1041.

Wardhaugh, Carl W., Stork, Nigel E., and Edwards, Will (2014) Canopy invertebrate community composition on rainforest trees: different microhabitats support very different invertebrate communities. Austral Ecology, 39 (4). pp. 367-377.

Grimbacher, Peter S., Nichols, Cassandra, Wardhaugh, Carl W., and Stork, Nigel E. (2014) Low host specificity of beetles associated with fruit falls in lowland tropical rainforest of north-east Australia. Austral Entomology, 53 (1). pp. 75-82.

Wardhaugh, Carl W. (2013) Estimation of biomass from body length and width for tropical rainforest canopy invertebrates. Australian Journal of Entomology, 52 (4). pp. 291-298.

Wardhaugh, Carl W., Edwards, Will, and Stork, Nigel E. (2013) Variation in beetle community structure across five microhabitats in Australian tropical rainforest trees. Insect Conservation and Diversity, 6 (4). pp. 463-472.

Wardhaugh, Carl W., Stork, Nigel E., and Edwards, Will (2013) Specialization of rainforest canopy beetles to host trees and microhabitats: not all specialists are leaf-feeding herbivores. Biological Journal of the Linnean Society, 109 (1). pp. 215-228.

Wardhaugh, Carl W., Edwards, Will, and Stork, Nigel E. (2013) Body size variation among invertebrates inhabiting different canopy microhabitat: flower visitors are smaller. Ecological Entomology, 38 (1). pp. 101-111.

Wardhaugh, Carl W., Stork, Nigel E., Edwards, Will, and Grimbacher, Peter S. (2013) Insects on flowers: the unexpectedly high biodiversity of flower-visiting beetles in a tropical rainforest canopy. Communicative & Integrative Biology, 6 (1). e22509. pp. 1-3.

Wardhaugh, Carl W. (2013) The importance of flowers for beetle biodiversity and abundance. In: Lowman, Margaret, Devy, Soubadra, and Ganesh, T., (eds.) Treetops at Risk: challenges of global canopy ecology and conservation. Springer, New York, NY, USA, pp. 275-288.

Wardhaugh, Carl W., Stork, Nigel E., and Edwards, Will (2012) Feeding guild structure of beetles on Australian tropical rainforest trees reflects microhabitat resource availability. Journal of Animal Ecology, 81 (5). pp. 1086-1094.

Wardhaugh, Carl W., Stork, Nigel E., Edwards, Will, and Grimbacher, Peter S. (2012) The overlooked biodiversity of flower-visiting invertebrates. PLoS ONE, 7 (9). e45796. pp. 1-8.

Wardhaugh, Carl William (2011) Microhabitat utilisation and the spatial distribution of rainforest canopy invertebrate communities. PhD thesis, James Cook University.